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We contacted the USGS Biological Resources Division and received a reply from Terry D'Erchia at the USGS Office of Biological Informatics and Outreach.
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Dear Madeline:

        Thank you for your interest in the USGS Biological Resources Division. We work with others to provide the scientific understanding and technologies needed to support the sound management and conservation of our Nation's biological resources.

        Because we are a Federal agency, the images on our Website - unless otherwise indicated - are in the public domain and as such are not copyrighted.  We do receive many requests for use of both the copyrighted and non copyrighted images that appear on our Web pages; permission is typically granted, with a request for attribution to the U.S. Department of the Interior, U.S. Geological Survey. In the case of images that we maintain through permission from another source, we ask that the requester go back to the original source to obtain permission for use.

I hope this response will be helpful for your project.  Good luck!

Sincerely,
Terry D'Erchia
USGS Office of Biological Informatics and Outreach