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Plants
Osmosis
and plant cells
Although
the osmotic principles apply equally to plant and animal cells,
a different set of terms is currently applied to the osmotic relationship
of plant cells. Water potential is a measure of the tendency of
water to leave a solution. Pure water is designated a water potential
of zero. As the solute molecules in a solution tend to prevent the
water molecules leaving it, the solution will have a lower water
potential than pure water. Its value will therefore be less than
zero, i.e. negative. The more concentrated the solution, the more
negative is its water potential.
For
practical purposes a plant cell can be considered as a solution
of salts and sugars in the vacuole surrounded by a partially permeable
membrane (tonoplast, cytoplasm and plasma membrane) and a slightly
elastic but completely permeable cell wall. A plant cell therefore
has a more negative water potential than pure water and will draw
in water when surrounded by it. This entry of water forces the living
part of the cell, known as the protoplast, against the cell wall.
In effect, the water in the vacuole is being subjected to a pressure
from the cell wall. This pressure is referred to as the pressure
potential. In a turgid plant cell this is a positive value, although
in the xylem of a transpiring plant it is negative. The water potential
of a cell is changed by the presence of solute molecules. This change
is referred to as the solute potential. As solute molecules invariably
lower the water potential, its value is always negative. The relationship
between these three terms is given as:
water
potential = solute potential + pressure potential
Water for the support of
plants
Water is almost impossible
to compress (press into a smaller volume). This feature of water
allows it to be used as a form of support by many smaller plants.
Although such plants do not have a skeleton, they do have cells
with strengthened walls the cell is filled with a gel-like substance
called cytoplasm. The cytoplasm contains a lot of water. Plant cells
also contain a large vacuole (a fluid-filled structure bound by
a membrane). The vacuole holds a lot of water. If the plant cell
takes up more water, the cytoplasm and the vacuole fill with water
and become swollen. This causes the contents of the cell to push
against the cell wall. These cells are described as being turgid,
and turgid cells make the plant stiffer and better able to stand
up to forces such as wind.
The importance of water
is clearly demonstrated by a plant that has not had enough water
because it soon starts to wilt. As the cell lose water, the plant
loses its support and the leaves begin to collapse. If the plant
is quickly given some water, the cell can recover. However, if a
plant is allowed to remain without water for too long, the cell
will be permanently damaged and the plant will die.
Water is a rare resource
in many parts of the world. Deserts are places where there is very
little water for almost all of the year. When it eventually does
rain, it usually comes all at once in heavy downpours, often causing
floods. In many other climatic areas of the world there are rainy
seasons and dry seasons. In these places, water has to be used carefully
and stored for use in the seasons.
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