Segregation

            In Mendel’s first experiment, he first established pure lines of tall and short garden pea plants, by allowing them to self-pollinate for several generations.  He then cross-pollinated these plants by first placing pollen from the short plants onto the stigmas of flowers on tall plants.  By examining the first generation (F1), he was able to determine the law of dominance.  He knew that all the plants in the F1 generation that were tall were hybrid for the traits he was looking for. 

            In Mendel’s first experiment, he first established pure lines of tall and short garden pea plants, by allowing them to self-pollinate for several generations.  He then cross-pollinated these plants by first placing pollen from the short plants onto the stigmas of flowers on tall plants.  By examining the first generation (F1), he was able to determine the law of dominance.  He knew that all the plants in the F1 generation that were tall were hybrid for the traits he was looking for.  Mendel next began to cross tall, hybrid plants, which appeared no different from pure tall plants, and view the results of that hybridization (the F2 generations).  The result of this experiment was that there were three times as many tall plants as there were short one.  This led Mendel to his Law of Segregation:  “When large numbers of hybrids are crossed, the factors segregate and then recombine to produce dominant and recessive offspring in a ration of 3:1.”  Because Mendel was before the discovery of DNA, he correctly attributed the transmission of traits to “factors,” which he could not describe, but did know their function.  Today, these factors would be known as alleles.

            It is much easier to understand the results of Mendel with the use of a Punnett square.  In order to create such a square to compare the possible results of one cross with respect to one trait, the two alleles donated by one partner are placed above a box, while the two alleles of the other are placed on the side.

                

T

T

t

Tt

Tt

t

Tt

Tt

            In the square above, the “T” represents the dominant tall gene, which creates a tall plant, while the “t” represents the recessive tall gene, or the phenotype short.  In the case of this cross, the phenotype would be all tall, as only tall plants would be created, while the genotype equals 100& hybrid.  These are the result obtained by Mendel in his first experiment.  When he crossed the hybrid plants to create the F2 generation, he created a cross like the one below and created the F2 generation.

           

T

t

T

TT

Tt

t

Tt

tt

           This time, the phenotype of the offspring is 75% tall, 25% short and the genotype of the offspring is 25% homozygous tall, 50% heterozygous tall, and 25% homozygous short.  Because the allele for a short plant is recessive, it will only appear when it is the only gene present. 

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