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Genetic Drift

No external factors are involved in genetic drift, which is a chance occurrence. However, a population of infinite size cannot occur Hardy-Weinberg Equilibrium so genetic drift is possible.

The size of the population is inversely proportionate to the gene frequency changes due to genetic drift and thus, a larger number of individuals reproducing results in small genetic drift effects. This relationship is similar in populations though the actual number of individuals is not the important factor here. "Effective" population size refers to the number of individuals producing offspring.

The effects of genetic drift are only pronounced over several generations, which consist of thousands of reproducing individuals and if the other process of evolution are missing, fixed populations would occur. Genetic drift causes an allele to have a precise frequency (e.g. an allele with frequency of 0.9 would definitely have a 90% chance of being present in the next generation).

An experiment conducted with eight individual fruit flies resulted in the discovery of the fact that as generations past, the allele frequencies will gradually decrease as is seen in this example (where the initial frequency was 0.5 and the final frequency was 0.23.)

Two types of genetic drift commonly observed include the founder effect and the bottleneck. The founder effect occurs when allele frequencies in a group of migrants differ from their population of origin. These individuals may develop distinct different traits if isolated. This results in species in neighbouring territories being more heterogeneous than species in adjacent territories.

Another type of special genetic drift is the bottleneck, which occurs when a dramatic decrease in population size occurs. A number of reasons including climatic, ecological changes and natural disasters could be responsible. A recovery of the original population size is possible though the frequencies of the alleles have been considerably altered, possibly affecting future generations of the species. An ideal condition for the occurrence of bottleneck is the large organism sizes due to the fact that typically such populations consist of fewer individuals. Persistent bottlenecks, however, may result in a decrease in the genetic variation thus altering the chance of future evolution of that species.