Movement
PatternsThe giant pandas of Wolong do not move about their home ranges
to any great extent. A resident, especially a young lower status or inexperienced animal,
will visit as little as 10 per cent of its home range in any one month, and some parts are
visited only rarely. This is atypical of carnivore behaviour, a feature of which is
extensive foraging movements within home ranges. The giant panda's strategy here is not
altogether surprising as the species is a carnivore only taxonomically. In practice, they
are herbivores. The only time they need to move extensively is during bamboo dieback, when
migration to other food areas is forced on them. Males have no favorite area, or core
area, and tend to use more of their range than females. Females concentrate their
activities within a core area (30-40 hectares, or 74-99 acres) which contains most of the
resources they need. A female will share her core area with adult males only, because
although she does not patrol the boundary of her core area, this zone is sacrosanct and
sub-adults and other females are regarded as trespassers.
Foraging movements depend on where the best available food is in any season. Spring is
an active time for giant pandas, when they make good use of their home ranges in search of
stands of umbrella bamboo. In spring 1981, a poor season for the shoots of this bamboo,
one of the radio-collared females, Zhen, actually crossed over Wolong Valley to the
opposite slope in search of food. Not only was this a long journey for a giant panda but a
very risky one as well, across a cultivated, populous valley lacking any cover. The
following two years were good food years and Zhen made no attempts to cross over again.
Summer is when pandas are at their laziest. They make many fewer excursions around
their home ranges and loll about eating long picnics of bamboo. Things hot up again in the
winter, a time of dwindling food supplies and, for males, the period leading up to finding
a mate. During this period, adult males use more than half of their home range and are
busy keeping tabs on the females within it. Significantly, sub-adult males, which are not
yet ready to breed, use less than half of their ranges during this period.
Female home ranges are located in the best areas. Best for food, for microclimate, for
nesting sites and for foraging. This translates into a number of habitat features: almost
continuous cover of arrow bamboo; south- and west-facing exposures, where snow does not
persist for as long as on other inclines; gently sloping ground, which avoids strenuous
climbing and saves precious energy; hollow trees or caves for dry, secure den sites; and a
well-wooded area with an almost continuous forest canopy. In open areas bamboo grows more
densely than beneath a canopy and the stalks are drier and tougher. They also tend to be
stunted and more leaves die in winter, which reduces the nutritional content. A good
canopy supports less dense, more nutritious bamboo groves and also gives shelter from the
rain and snow. The large trees that form the canopy also provide den sites.
The main imperative for a male is to maximize access to potential mates, which means
home range overlap with as many females as possible. Sub-adults draw the short straw in
this situation. They have to make do with less than ideal living quarters. In the Wolong
study two sub-adults that had formed a loose acquaintance, Long and Ning, shared a home
range on a steep hillside cut by many ravines and almost denuded of trees. They
concentrated their foraging in and around the remaining forest patches rather than
attempting to exploit the open slopes. This shows that open habitat is clearly not
preferred and implies a level of competition between adults and sub-adults for resources.
By carnivore standards, giant pandas are modest landowners. Home ranges are small,
varying from 3.9 to 6.4 km square, and female ranges are almost the same size as those of
their male neighbours. This is quite unlike the situation in large carnivores such as the
black bear and grizzly bear, where male home ranges may be several times larger than those
of females and where males particularly frequent all parts of their home ranges. The
reason for this difference, no doubt, lies once again in the giant panda's limited menu.
It may be that there is a limit to the size of land that a male can monitor on his
low-nutrient diet. He is constrained to spend most of his time searching for, and eating,
bamboo. As with most other mammals, the size of a giant panda's home range changes as the
individual matures. Young pandas start off by sharing their mother's home range. Then,
after they part company with her, they may make exploratory trips into new areas,
sometimes travelling as much as 14 km (8.5 miles) in a day (the norm is usually 2 km (1.2
miles) a day) to look for the best available area. Once the choice has been made and they
are ready to mate, the young adult will settle down into a home range of its own.